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When you create a new product from BCC, you can define parent categories in categorization tab.
To create sub categories of a category, add new category from categorization section in general tab of category.
From ACC you can do it through defaultParentCategory/fixedRelatedCategories which you will get while creating a new category.
Ancestor Categories will be populated automatically.
When you create a category under a pre existing category , category x under y say then y gets added as parent category , ancestor categories are computation based and when you run the Catalog Administration Service these get calculated and set by ATG internally you need not set them ( cannot set them using ACC).
The below links might help you in understanding