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"NEVER" transaction participation in the service adapter in exposed services lane means that no transaction context will be imported when this service will be called.
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The properties refer to different transaction support "contexts", as you can see in the docs bellow... The bpel.config.transaction refers to BPEL Process Manager Transaction Semantics, while the "Transaction Participation" refers to WS-AT which provides transaction interoperability between Oracle WebLogic Server and other vendors' transaction services... I've never tested myself but I'd guess if you are transacting among BPEL processes, the bpel.config.transaction will prevail...