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APEX resides in the database, so what you're possibly asking is can your HTTP server reside in Windows, connecting to the APEX instance on the database that is installed on Linux.
Yes you can.
Use database link (see http://docs.oracle.com/cd/B12037_01/server.101/b10759/statements_5005.htm) to read the DB in linux server. But as scott said, a Oracle DB must be in Windows to hold APEX and use DB link to target DB to connect.
You must ensure ports are opened and accessable from Windows DB, network ACL enabled (I think I got ACL related error last time I tried it)..etc.